Code:
\documentclass[12pt]{article}
\usepackage[ngerman, english]{babel}
\usepackage{lmodern}
\usepackage{geometry}
\geometry{margin=2cm,}
\usepackage{array}
\usepackage{booktabs}
\usepackage{caption}
\usepackage{siunitx}
\usepackage{pdflscape}
\usepackage{ragged2e}
\usepackage[para]{threeparttable} %Tabellen mit Legenden
\newcommand{\species}[1]{\textit{#1}}
\newcommand{\tabhead}[1]{\RaggedRight\bfseries #1\arraybackslash}
\begin{document}
\begin{landscape}
\begin{table}
\caption{Frequency of colonization of the root segments by different PAC species in 2004 and 2014}
\centering
\label{tab:Dyn2}
\begin{tabular}{lSSSSSSSSSS}
\toprule
& \multicolumn{2}{p{3.5cm}}{\RaggedRight 2004 ($n=5$ root segments per grid point)}
& \multicolumn{2}{p{3.5cm}}{\RaggedRight 2014 ($n=5$ root segments per grid point)}
& \multicolumn{2}{p{3.5cm}}{\RaggedRight Difference between years (z-test)}
& \multicolumn{2}{p{3.5cm}}{\RaggedRight Extended sampling 2014 ($n=30$ root segments per grid point}
& \multicolumn{2}{p{3.5cm}}{\RaggedRight Difference between normal and extended sampling in 2014 (z-test)} \\
\midrule
\tabhead{Species}
& \parbox{2.5cm}{\tabhead{No. of colonized root segments}}
& {\tabhead{\%}}
& \parbox{2.5cm}{\tabhead{No. of colonized root segments}}
& {\tabhead{\%}}
& {\tabhead{z value}}
& {\tabhead{p value}}
& \parbox{2.5cm}{\tabhead{No. of colonized root segments}}
& {\tabhead{\%}}
& {\tabhead{z value}}
& {\tabhead{p value}} \\
\midrule
\species{A. applanata} & 0 & 0 & 0 & 0 & {n.a.} & {n.a.} & 2 & 0.4 & {n.a.} & {n.a.} \\
\species{P. europaea} & 30 & 37.5 & 12 & 15.0 & 2.777 & 0.003** & 61 & 12.7 & -0.435 & 0.668 \\
\species{P. helvetica} & 9 & 11.3 & 11 & 13.8 & 0.447 & 0.327 & 40 & 8.3 & -1.153 & 0.876 \\
\species{P. letzii} & 5 & 6.3 & 7 & 8.8 & 0.577 & 0.282 & 29 & 6.0 & -0.704 & 0.759 \\
\species{P. turicensis} & 16 & 20.0 & 18 & 22.5 & 0.343 & 0.366 & 86 & 17.9 & -0.721 & 0.765 \\
\species{P. uotilensis} & 1 & 1.3 & 0 & 0 & {n.a.} & {n.a.} & 1 & 0.2 & {n.a.} & {n.a.} \\
\bottomrule
\end{tabular}
\end{table}
\end{landscape}
\end{document}
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